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Mules absolutely do require fuel; they’ll only be useful for a few days without food.
Additionally, they require a lot of rest and need to be rotated out more frequently than humans. Nazi Germany was very dependent on draft animals on WWII and the logistics were nuts.
Any type of grass or non poisonous vegetation works.
In difficult terrain animals can still be the best choice for transport. Dont know if that applies here, but i know from Germanys invasion of Ukraine in WW2 that the mud in Spring and Autumn is terrible to traverse with mechanized units.
Not so easy during winter.
You can’t just leave them grazing wherever. Mines and artillery kill them just as well.
Some fodder will need to be brought in for them.